A pod of killer whales swims



Asper, E. D., G. W. Young and M. T. Walsh. 1988. Observations on the Birth and Development of a Captive born Killer Whale. International Zoo Yearbook 27. Pp. 295-304.

Au, W. W. L. 1993. The Sonar of Dolphins. New York: Springer-Verlag Inc.Au, W. W. L. 2009. Echolocation. In: Perrin W. F., B. Wursig and J. G. M. Thewissen. Encyclopedia of Marine Mammals, 2nd ED, San Diego, CA: Academic Press: pp. 348-357.

Au, W. W. L., J. K. B. Ford, J. K. Horne and K. A. Newman Allman. 2004. Echolocation signals of free-ranging killer whales (Orcinus orca) and modeling of foraging for Chinook salmon (Oncorhynchus tshawytscha). Journal of the Acoustical Society of America. 115(2): 901-909.

Ayres, K. L., R. K. Booth, J. A. Hempelmann, K. L. Koski, C. K. Emmons, R. W. Baird, K. Balcomb-Bartok, M. B. Hanson, M. J. Ford and S. K. Wasser. 2012. Distinguishing the Impacts of Inadequate Prey and Vessel Traffic on an Endangered Killer Whale (Orcinus orca) Population. PLOS One. 7(6): 1-12.

Bain, D. 1990. Examining the validity of inferences drawn from photo-identification data, with special reference to studies of the killer whale (Orcinus orca) in British Columbia. Reports of the International Whaling Commission Special Issue 10: 93-100.

Baird, R. W. 2000. The Killer Whale: Foraging Specializations and Group Hunting. In: J. Mann, R. C. Connor, P. L. Tyack and H. Whitehead, editors. Cetacean societies: Field Studies of Dolphins and Whales. University of Chicago Press, Chicago, Illinois. Pp. 127-153.

Baird, R. W., Lawrence M. Dill and M. Bradley Hanson. 1998. Diving Behaviour of Killer Whales. Abstracts of the World Marine Mammal Science Conference, Monaco. Pg. 8.

Baird, R. W., M. B. Hanson, E. A. Ashe, M. R. Heithaus and G. J. Marshall. 2003. Studies of foraging in "southern resident" killer whales during July 2002: dive depths, bursts in speed, and the use of a "Crittercam" system for examining subsurface behavior. National Marine Fisheries Service, National Marine Mammal Laboratory, Seattle, Washington.

Baird, R. W., D. McSweeney, C. Bane, J. Barlow, D. R. Salden, L. R. K. Antoine, R. G. LeDuc and D. L. Webster. 2006. Killer whales in Hawaiian waters: Information on population identity and feeding habits. Pacific Science. 60(4): 523-530.

Barnes, L. G. 1990. The Fossil Record and Evolutionary Relationships of the Genus Tursiops. In: Leatherwood, S. and Reeves, R.R., eds., The Bottlenose Dolphin. New York: Academic Press. pp. 3-26.

Barrett-Lennard, L.G., J. K. B. Ford and K. A. Heise. 1996. The mixed blessing of echolocation: differences in sonar use by fish-eating and mammal-eating killer whales. Animal Behavior. 51:553-565.

Benirschke, K., and L. H. Cornell. 1987. The placenta of the killer whale, Orcinus orca. Marine Mammal Science 3:82–86.

Berta, A., J. L. Sumich and K.M. Kovacs. 2006. Marine Mammals: Evolutionary Biology,2ndEd. Academic Press, Burlington, MA. Pp. 105.

Bigg, M. A. 1982. An Assessment of Killer Whale (Orcinus orca) Stocks off Vancouver Island, British Columbia. Rep. Int. Whal. Commn. 32. Pp. 655-666.

Bigg, M. A., G. M. Ellis, J. K. B. Ford and K. C. Balcomb. 1987. Killer Whales-a Study of Their Identification, Genealogy and Natural History in British Columbia and Washington State. Phantom Press, Nanaimo.

Black, N. A., A. Schulman-Janiger, R. L. Ternullo and M. Guerrero-Ruiz. 1997. Killer whales of California and western Mexico: A Catalog of photo-identified individuals. U.S. Dep. Commer., NOAA Tech. Memo. NMFSS¬WFSC-247. 174p.

Bowers, C. A. and R.S. Henderson. 1972. PROJECT DEEP OPS: Deep Object Recovery with Pilot and Killer Whales. NUC TP 306. Undersea Surveillance and Ocean Sciences Department, Naval Undersea Center, San Diego, CA. Unclassified.

Bowles, A. E., W. G. Young and E. D. Asper. 1988. Ontogeny of Stereotyped Calling of a Killer Whale Calf, Orcinus orca, During Her First Year. In: J. Sigurjonsson and S. Leatherwood (eds.), North Atlantic Killer Whales, Rit Fiskideildar 11. Reykjavik. Pp. 251-275.

Boyd, I. L., C. Lockyer and H. D. Marsh. 1999. Reproduction in Marine Mammals. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press. Washington. Pp. 218-286.

de Bruyn P. J., C. A. Tosh and A. Terauds. 2013. Killer whale ecotypes: is there a global model? Biol Rev Camb Philos Soc. Feb;88(1):62-80.

Carwardine, Mark, Erich Hoyt, R. Ewan Fordyce and Peter Gill. 1998. Whales, Dolphins & Porpoises. The Nature Company Guides. New York: Dorling Kindersley, Inc.

Clark, R. B. 1997. Marine pollution. 4th edition. Clarendon Press, Oxford, United Kingdom.

Clark, S. T., and D. K. Odell. 1999a. Nursing parameters in captive killer whales (Orcinus orca). Zoo Biology 18:373–384.

Clark, S. T., and D. K. Odell 1999b. Allometric relationships and sexual dimorphism in captive killer whales (Orcinus orca). Journal of Mammalogy 80:777–785.

Clark, Steven T., Daniel K. Odell and C. Thad Lacinak. 2000. Aspects of Growth in Captive Killer Whales (Orcinus orca). Marine Mammal Science, Volume 16, Issue 1: 110–123.

Christensen, Ivar. 1984. Growth and Reproduction of Killer Whales, Orcinus orca, in Norwegian Coastal Waters. Report of the International Whaling Commission. Special Issue 6: pp. 253-258.

Committee on Taxonomy. 2012. List of marine mammal species and subspecies. Society for Marine Mammalogy, www.marinemammalscience.org, consulted on May 12, 2014.

Couperus, A. S. 1994. Killer whales (Orcinus orca) scavenging on discards of freezer trawlers north east of Shetland islands. Aquatic Animals, 20.1: pp. 47-51.

Cornell. L. H. 1983. Hematology and clinical chemistry values in the killer whale (Orcinus orca). Journal of Wildlife Diseases 19:259–264.

Crance, J. L., A. E. Bowles and A. Garver. 2013. Evidence for vocal learning in juvenile male killer whales, Orcinus orca, from an adventitious cross-socializing experiment. Journal of Experimental Biology 217:1229–1237.

Cutnell, John D. and Kenneth W. Johnson. 1998. Physics. 4th ed. New York: Wiley. Pg. 466.

Dahlheim, M. E. and F. Awbrey. 1982. A classification and comparison of vocalizations of captive killer whales (Orcinus orca). Journal of the Acoustic Society of America 72:661–670.

Dahlheim, M., A. Schulman-Janiger, N. Black, R. Ternullo, D. Ellifrit and K. Balcomb III. 2008. Eastern temperate North Pacific offshore killer whales (Orcinus orca): Occurrence, movements, and insights into feeding ecology. U.S. Department of Commerce: Publications, Agencies, and Staff of the U.S. Department of Commerce.

Dahlheim, Marilyn Elayne. 1988. Killer Whale (Orcinus orca) depredation on longline catches of sablefish (Anoplopoma fimbria) in Alaskan waters. Seattle: NWAFC Processed Report, National Marine Fisheries Service.

Deecke, V. B., J. K. B. Ford, and P. J. B. Slater. 2005. The vocal behaviour of mammal-eating killer whales: communicating with costly calls. Animal Behaviour 69: pp. 395-405.

Duffield, Deborah A. and K. W. Miller. 1988. Demographic Features of Killer Whales in Oceanaria in the United States and Canada,1965-1987. Rit Fiskideildar 11: pp. 297-306.

Duffield, Deborah A., Daniel K. Odell, James F. McBain and Brad Andrews. 1995. Killer Whale Reproduction at Sea World. Zoo Biology 14: pp. 417-430.

Duffus, David A. and Philip Dearden. 1993. Recreational Use, Valuation, and Management of Killer Whales (Orcinus orca) on Canada's Pacific Coast. Environmental Conservation, Vol. 20 (2).

Durban, J. and V. Deeke. 2011. How do we study killer whales? Whalewatcher Killer Whale: The Top, Top Predator. In Pitman, R. L. (Ed.) Journal of the American Cetacean Society. 40(1). 2011.

Durban, J. W. and R. L. Pitman. Antarctic killer whales make rapid, round-trip movements to subtropical waters: evidence for physiological maintenance migrations? Biology Letters. 8:274-277. 2011.

Elsner, R. 1999. Living in Water: Solutions to Physiological Problems. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press.Washington: pp. 73-116.

Ellis, Richard. 1989. Dolphins and Porpoises. New York: Alfred A. Knopf, Inc: pp. 167-189.

Fisheries and Oceans Canada. 2007. Recovery Strategy for the Transient Killer Whale (Orcinus orca) in Canada. Species at Risk Act Recovery Strategy Series. Fisheries and Oceans Canada, Vancouver, vi + 46 pp.

Foote, A. 2011. North Atlantic Killer Whales. In Whalewatcher Killer Whale: The Top, Top Predator. Pitman, R. L. (Ed.) Journal of the American Cetacean Society. 40(1): 30-32.

Foote, A. D., J. Newton, S. B. Piertney, E. Willerslev, and M. T. P. Gilbert. 2009. Ecological, Morphological and Genetic Divergence of Sympatric North Atlantic Killer Whale Populations. Molecular Ecology. 18(24): pp. 5207-5217.

Ford, J. K. B., G. Ellis and K. C. Balcomb. 1994. Killer Whales: the Natural History and Genealogy of Orcinus orca in British Columbia and Washington State. Canada: UBC Press.

Ford, J. K. B., G. M. Ellis, L. G. Barrett-Lennard, A. B. Morton, R. S. Palm and K. C. Balcomb III. 1998. Dietary specialization in two sympatric populations of killer whales (Orcinus orca) in coastal British Columbia and adjacent waters. Canadian Journal of Zoology 76: pp. 1456-1471.

Ford, J. K. B., G. M. Ellis and K. C. Balcomb. 2000. Killer Whales: the Natural History and Genealogy of Orcinus orca in British Columbia and Washington State. 2nd ed. UBC Press, Vancouver, British Columbia.

Ford, J. K. B., G. M. Ellis, C. O. Matkin, M. H. Wetklo, L. G. Barrett-Lennard and R. E. Wither. 2011. Shark Predation and Tooth Wear in a Population of Northeastern Pacific Killer Whales. Aquatic Biology. 11: pp. 213-224.

Ford J. K. B. 2002. Killer Whale Orcinus orca. In: Perrin W. F., B. Wursig and J. G. M. Thewissen. Encyclopedia of Marine Mammals, San Diego, CA: Academic Press: pp. 669-676.

Ford J. K. B. 2009. Killer Whale Orcinus orca. In: Perrin W. F., B. Wursig and J. G. M. Thewissen. Encyclopedia of Marine Mammals, 2nd edn, San Diego, CA: Academic Press: pp. 650-657.

Forney, K. A. and P. Wade. 2006. Worldwide Distribution and Abundance of Killer Whales. In: J. A. Estes, R. L. Brownell, Jr., D. P. DeMaster, D. F. Doak and T. M. Williams (editors). Whales, Whaling and Ocean Ecosystems. University of California Press, Berkeley, California: pp. 145-162.

Fraser, F. C. 1949. Whales and Dolphins. Field Book of Giant Fishes. Eds J. R. Norman and F. C. Fraser. Pp. 201-349.G. Putman's Son, New York.

Friedl, W. A., P. E. Nachtigall, P. W. B. Moore, N. K. W. Chun, J. E. Haun, and R. W. Hall. 1990. Taste Reception in the Pacific Bottlenose Dolphin (Tursiops truncatus gilli) and the California Sea Lion (Zalophus californianus.) In: J.A. Thomas and R.A. Kastelein. (eds.) Sensory Abilities of Cetaceans: Laboratory and FieldEvidence. Series A: Life Sciences Vol. 196. Plenum Press. New York, NY.: pp. 447-454.

Funke, C., D. P. King, J. F. McBain, D. Adelung and J. L. Stott. 2003. Expression and functional characterization of killer whale (Orcinus orca) interleukin-6 (IL-6) and development of a competitive immunoassay. Veterinary Immunology and Immunopathology 93:69–79.

Graham, Mark S. and Pierre R. Dow. 1990. Dental Care for a Captive Killer Whale, Orcinus orca. Zoo Biology Vol. 9 (4): pp. 325-330.

Grant, S. C. H. and P. S. Ross. 2002. Southern resident killer whales at risk: toxic chemicals in the British Columbia and Washington environmentIllinois. Pp. 127-153. Canadian Technical Report of Fisheries and Aquatic Sciences 2412: pp. 1-111.

Grolier Publishing. 1967. "Body, Human". The New Book of Knowledge. New York: Grolier: pp. 285.

Guinet, Christophe. 1990. Intentional Stranding in Killer Whales. Whalewatcher. San Pedro: The American Cetacean Society, Volume 24 (3): pp. 9-11.

Haenel, N. J. 1986. General notes on the behavioral ontogeny of Puget Sound killer whales and the occurrence of allomaternal behavior. In: B. C. Kirkevold and J. S. Lockard, editors. Behavioral Biology of Killer Whales.Alan R. Liss, New York, New York: pp. 285-300.

Hall, J. D., and C. S. Johnson. 1972. Auditory thresholds of a killer whale Orcinus orcaLinnaeus. Journal of the Acoustic Society of America 51:515.

Hammers, A. M. 2003. Killer Whales Killing Other Whales. National Geographic Today.

Hanson, M. B., R. W. Baird, J.K.B. Ford, J. Hempelmann-Halos, D. M. Van Doornik, J. R. Candy, C. K. Emmons, G. S. Schorr, B. Gisborne, K. L. Ayres, S. K. Wasser, K. C. Balcomb, K. Balcomb-Bartok, J. G. Sneva and M. J. Ford. 2010. Species and stock identification of prey consumed by endangered southern resident killer whales in their summer range. Endangered Species Research. 11:69-82.

Hedrick, M. S., and D. A. Duffield. 1991. Hematological and rheological characteristics of blood in seven marine mammal species: physiological implications for diving behaviour. Journal of Zoology (London) Herman, Louis M., ed. 1980. Cetacean Behavior: Mechanics and Functions. New York: John Wiley and Sons. 225:273–283.

Herman, Louis M., ed. 1980. Cetacean Behavior: Mechanics and Functions. New York: John Wiley and Sons.

Heyning, John E. 1988. Presence of Solid Food in a Young Killer Calf Whale (Orcinus orca). Marine Mammal Science 4: pp. 68-71.

Heyning, John E. and Marilyn E. Dahlheim. 1999. Killer Whale-Orcinus orca. Handbook of Marine Mammals, Vol. 6, 11: pp. 281-322.

Heyning, John E. and Marilyn E. Dahlheim. 1988. Orcinus orca. Mammalian Species. American Society of Mammalogists.

Hickie, B. E., P. S. Ross, R. W. Macdonald and J. K. B Ford. 2007. Killer whales (Orcinus orca) face protracted health risks associated with lifetime exposures to PCBs. Environmental Science & Technology 41: pp. 6613-6619.

Hoyt, E. 2001. Whale watching 2001: worldwide tourism numbers, expenditures, and expanding socioeconomic benefits. International Fund for Animal Welfare, Yarmouth, Massachusetts.

Hoyt, E. 1990. Orca: the Whale Called Killer. 3rd edition. Camden House Publishing, North York, Ontario.

Jacobs, Fred. 2013. http://www.cnn.com/2013/10/21/us/seaworld-blackfish-qa/ consulted on May 2, 2014.

Jacobsen, Jeff. 1990. The Social Ways of Sleeping Orca. Whalewatcher. San Pedro: The American Cetacean Society, Volume 24 (3): pp. 6-8.

Jefferson, T. A., P. J. Stacey, and R. W. Baird. 1991. A Review of Killer Whale Interactions with Other Marine Mammals: Predation to Co-existence. Mammal Reviews, 21 (4): pp. 151-180.

Jefferson, T. A., Leatherwood, Stephen and Marc A. Webber. 1993. FAO Species Identification Guide. Marine Mammals of the World. Rome: Food and Agriculture Organization of the United States.

Jefferson, T. A., Stacey, P. J. & Baird, R. W. 1991. A review of killer whale interactions with other marine mammals: Predation to co-existence. Mammal Review 21 (4): pp. 151-180. Killer Whales in the Eastern North Pacific (Orcinus orca). 2002. Marine Mammal Commission-Annual Report for 2002: pp. 51-57.

Jefferson, T. A., Webber, M. A., and R. L. Pitman. 2008. Marine Mammals of the World: A Comprehensive Guide to their Identification. Amsterdam, Academic Press. Pp. 158-163.

Kasamatsu, F. and G. G. Joyce. 1995. Current status of odontocetes in the Antarctic. Antarctic Sci 7: 365-379.

Kastelein, R.A., S. Walton, D. Odell, S. H. Nieuwstraten, and P. R. Wiepkema. 2000. Food consumption of a captive female killer whale (Orcinus orca). Aquatic Mammals 26:127–131.

King, D. P., et al. 1996. Molecular cloning and sequencing of interleukin DNA fragments from the harbor seal (Phoca vitulina), killer whale (Orcinus orca), and Southern sea otter (Enhydra lutris nereis). Immunogenetics 43:190–195.

Kooyman, G.L. Diving physiology. In Encyclopedia of Marine Mammals, 2nd Edition. Perrin, W. F., B. Würsig and J. G. M. Thewissen. Burlington, MA. Associated Press. 2009.

Krahn, M. M., M. J. Ford, W. F. Perrin, P. R. Wade and R. P. Angliss. 2004. Status review of southern resident killer whales (Orcinus orca) under the Endangered Species Act. NOAA Technical Memo NMFS-NWFSC-62. Northwest Fisheries Science Center, Seattle, WA.

Krahn, M. M., M. B. Hanson, G. S. Schorr, C. K. Emmons, D. G. Burrows, J. L. Bolton, R. W. Baird and G. M. Ylitalo. 2009. Effects of age, sex and reproductive status on persistent organic pollutant concentrations in "Southern Resident" killer whales. Marine Pollution Bulletin. 58: pp. 1522-1529.

Kuczaj, S. (ed.) 2010a. Research with Captive Marine Mammals is Important Part I. International Journal of Comparative Psychology 23(3): pp. 225-534.

Kuczaj, S. (ed.) 2010b. Research with Captive Marine Mammals is Important Part II. International Journal of Comparative Psychology 23(4): pp. 536-825.

Lane, Sally Jean. 1996. "Associations and the Ontogeny of Calling in a Killer Whale Calf, Orcinus orca, During Her First Six Months." M. S. dissertation, University of San Diego.

Levenson, D. H. and A. Dizon. 2003. Genetic Evidence for the Ancestral Loss of Short-Wavelength-Sensitive Cone Pigments in Mysticete and Odontocete Cetaceans. Proceedings: Biological Sciences: pp. 673-679.

Lyamin, O., P. R. Manger, S. H. Ridgway, L. M. Mukhavetov and J. M. Siegel. 2008. Cetacean sleep: An unusual form of cetacean sleep. Neuroscience and Biobehavioral Reviews 32: pp. 1451-1484.

Lyamin, O., L. M. Mukhametov and J. M. Siegel. 2004. Relationship Between Sleep and Eye State in Cetaceans and Pinnipeds. Archives Italiennes de Biologie, 142: pp. 557-568.

Lyamin, O., J. Pryaslova, V. Lance and J. M. Siegel. 2005. Continuous activity in cetaceans after birth. Nature, 435: pg. 1177.

Mass, A. M. and A. Ya. Supin. 2007. Adaptive features of aquatic mammals' eye. The Anatomical Record. 290: 701-715.

Matkin, C. O., E. L. Saulitis, G. M. Ellis, P. Olesuik and S. D. Rice. 2008. Ongoing population-level impacts on killer whales Orcinus orca following the 'Exxon Valdez' oil spill in Prince William Sound, Alaska. MarineEcology Progress Series. 356: pp. 269-281.

Matkin, C. O. and E. L. Saulitis. 1997. Restoration Notebook: Killer Whale (Orcinus orca). Exxon Valdez Oil Spill Trustee Council, Anchorage, Alaska.

Matkin, C.O. and E. L. Saulitis. 1994. Killer whale (Orcinus orca) Biology and Management in Alaska. Contract Number T75135023. Marine Mammal Commission, Washington D.C. 46 pp.

Matkin, C. O., John W. Durban, Eva L. Saulitis, Russel D. Andrews, Janice M. Straley, Dena R. Matkin and Graeme M. Ellis. 2012. Contrasting abundance and residency patterns of two sympatric populations of transient killer whales (Orcinus orca) in the northern Gulf of Alaska. Fish Bulletin. 110(2):143-155.

Matkin, C.O., J. Ward Testa, Graeme M. Ellis and Eva L. Saulitis. 2013. Life history and population dynamics of southern Alaska resident killer whales (Orcinus orca). Marine Mammal Science: 10.1111/mms.12049.

Miller, L. J. 2009. The Effects of Dolphin Education Programs on Visitors' Conservation-Related Knowledge, Attitude and Behavior. PhD Dissertation. University of Southern Mississippi, Hattiesburg, MS. 62 pages.

Mitchell, Edward and Alan N. Baker. 1980. Age of reputedly old killer whale, Orcinus orca, 'Old Tom' from Eden, Twofold Bay, Australia. Rep. International Whaling Commission, Special Issue 3.

Moore, S. E., J. K. Francine, A. E. Bowles and J. K. F. Ford. 1988. Ontogeny of Calls of Killer Whales Orcinus orca, from Iceland and Norway. In: J. Sigurjonsson and S. Leatherwood (eds.), North Atlantic Killer Whales,Rit Fiskideildar 11. Reykjavik: pp. 225-250.

Morin, P. A., F. I. Archer, and A. D. Foote. 2010. Complete mitochondrial genome phylogeographic analysis of killer whales (Orcinus orca) indicates multiple species. Genome Research. http://www.genome.org/cgi/doi/10.1101/gr.102954.109.

Nishiwaki, M. 1972. General Biology. In: S. H. Ridgway, ed., Mammals of the Sea: Biology and Medicine: pp. 3-204.

Nishiwaki, Masaharu and Chikao Handa. 1958. Killer Whales Caught in the Coastal Waters off Japan for Recent 10 Years. The Scientific Reports of the Whale Research Institute, 13: pp. 85-96.

National Marine Fisheries Service (NMFS). 2008. Recovery Plan for Southern Resident Killer Whales (Orcinus orca). National Marine Fisheries Service, Northwest Region, Seattle, Washington.

Nature.Learning in the wild: Much of what people know about science is learned informally. Education policy-makers should take note. 8 April 2010.464:813-814.

North Atlantic Killer Whale ID Project. http://www.northatlantickillerwhales.com accessed June 25, 2014.Nowak, R. M. and J. L. Paradiso. 1999. Killer Whale. Walker's Mammals of the World. Baltimore: The Johns Hopkins University Press, pp. 942-944.

Olesiuk, P.F. 2012. Population biology of the resident ecotype of killer whale in British Columbia. In Materials of the Killer Whale Workshop VII International Conference: Martine Mammals of the Holarctic.Suzdal, Russia. Pp. 3-8.

Olesiuk, P. F., M. A. Bigg and G. M. Ellis. 1990. Life history and population dynamics of resident killer whales (Orcinus orca) in the coastal waters of British Columbia and Washington State. Report of theInternational Whaling Commission, Special Issue 12: pp. 209-243.

Olson, P. A. and T. Gerrodette. 2008. Killer whales of the eastern tropical Pacific: A catalog of photo-identified individuals. NOAA Technical Memorandum. NOAA-NMFS-SWFSC-428, 120 p.

O'Shea, Thomas J. 1999. Environmental Contaminants and Marine Mammals. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press. Washington: pp. 485-564.

Pabst, Ann D., Rommel, Sentiel A. and William A. McLellan. 1999. The Functional Morphology of Marine Mammals. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press.Washington: pp. 15-72.

Paulos, Robin D., Marie Trone and Stan A Kuczaj II. 2010. Play in Wild and Captive Cetaceans. International Journal of Comparative Psychology, 23, 701-722.

Perrin, W. F. and S. B. Reilly. 1984. Reproductive Parameters of Dolphins and Small Whales of the Family Delphinidae. Report of the International Whaling Commission, Special Issue 6: pp. 97-134.

Pitman, R. L. (Ed.) 2011. Whalewatcher  Killer Whale: The Top, Top Predator. Journal of the American Cetacean Society. 40(1).

Pitman, R. L., J. W. Durban, M. Greenfelder, C. Guinet, M. Jorgensen, P. A. Olson, J. Plana, P. Tixier and J. R. Towers. 2011. Observations of a Distinct Morphotype of Killer Whale (Orcinus orca), Type D, from Subantarctic Waters. Polar Biology. 34: pp. 303-306.

Pitman, R. L. and J. W. Durban. 2011. Cooperative hunting behavior, prey selectivity and prey handling by pack ice killer whales (Orcinus orca), type B, in Antarctic Peninsula waters. Marine Mammal Science. 2011.

Pitman, R. L. and J. W. Durban. 2010. Killer whale predation on penguins in Antarctica. Polar Biology. 33: pp. 1589-1594.

Pitman, R. L., D. L. Perryman and E. Eilers. 2007. A Dwarf Form of Killer Whale in Antarctica. Journal of Mammalogy. 88(1): pp. 43-48.

Pitman, R. L. and P. Ensor. 2003. Three Forms of Killer Whales (Orcinus orca) in Antarctic Waters. Journal of Cetacean Research and Management. 5(2): pp. 131-139.

Pyle, P., M. J. Schramm, C. Keiper and S.D. Anderson. 1999. Predation on a White Shark by a Killer Whale and a Possible Case of Competitive Displacement. Marine Mammal Science. 5(2) April: pp. 563-568.

Reidarson, T. H., D. Duffield, and J. McBain. 2000. Normal hematology of marine mammals. Pp. 1164–1173 in Schalm's veterinary hematology (B. V. Feldman, J. G. Zinkl, and N. C. Jain, eds.). Lippincott Williams & Wilkins, Baltimore, Maryland.

Renner, M. and K. Bell. 2008. A white killer whale in the central Aleutians Arctic 61: 102-104.

Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press. Washington. 1999.

Ridgway, S. H. 2002. Asymmetry and Symmetry in Brain Waves from Dolphin Left and Right Hemispheres: Some Observations after Anesthesia, During Quiescent Hanging Behavior, and During Visual Obstruction Brain, Behavior and Evolution 60: pp. 265-274.

Ridgway, S. H. 1999. The Cetacean Central Nervous System. In: Adelman, G. and Smith, B.H., (eds.), Elsevier's Encyclopedia of Neuroscience, 2nd ed. Cambridge, Massachusetts: Elsevier Science Publishing Co.: pp. 352-358.

Ridgway, S. H. and D. A. Carder. 1990. Tactile Sensitivity, Somatosensory Responses, Skin Vibrations, and the Skin Surface Ridges of the Bottlenose Dolphin, Tursiops truncatus. In: Thomas, J.A., and Kastelein, R.A.,(eds.), Sensory Abilities of Cetaceans: Laboratory and Field Evidence. NATO ASI Series, Vol. 196. New York:Plenum Publishing: pp. 163-179.

Riesch, R. and V. B. Deecke. 2011. Whistle communication in mammal-eating killer whales (Orcinus orca): further evidence for acoustic divergence between ecotypes. Behavioral Ecology and Sociobiology 65: 1377-1387.

Robeck, T. R., and N. H. Nollens. 2013. Hematologic and serum biochemical parameters reflect physiological changes during gestation and lactation in killer whales (Orcinus orca). Zoo Biology 32:497–509.

Robeck, T. R., S. A. Gearhart, K. J. Steinman, E. Katsumata, J. D. Loureiro and J. K. O'Brien. 2011. In vitro sperm characterization and development of a sperm cryopreservation method using directional solidification in the killer whale (Orcinus orca). Theriogenology 76:267-279.

Robeck, T. R. and S. L. Monfort. 2006. Characterization of male killer whale (Orcinus orca) sexual maturation and reproductive seasonality. Theriogenology, 66(2): 242-250.Robeck, T.R., O'Brien, J.K. and D.K. Odell. Captive Breeding. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press. Washington: pp.178-183.

Robeck, T. R., A. L. Schneyer, J. F. McBain, L. M. Dalton, M. T. Walsh, N. Czekala and D. C. Kraemer, 1993. Analysis of urinary immunoreactive steroid metabolites and gonadotropins for characterization of the estrous cycle, breeding period, and seasonal estrous activity of captive killer whales (Orcinus orca). Zoo Biology, 12, 2: pp. 173-188.

Robeck, T. R., K. J. Steinman, S. Gearhart, T. R. Reidarson, J. F. McBain and S. L. Monfort. 2004. Reproductive physiology and development of artificial insemination technology in killer whales (Orcinus orca). Biology of Reproduction 71: 650-660.

Robeck, T. R., K. Willis, M. R. Scarpuzzi and J. K. O'Brien. 2015. Comparisons of life-history parameters between free-ranging and captive killer whale (Orcinus orca) populations for application toward species management. Journal of Mammalogy Advance Access published July 10, 2015.

Rommel, Sentiel. 1990. Osteology of the Bottlenose Dolphin. In Leatherwood, Stephen and Randall R. Reeves, The Bottlenose Dolphin. San Diego: Academic Press, Inc.: pp. 29-50.

Ross, P. S., G. M. Ellis, M. G. Ikonomou, L. G. Barrett-Lennard and R. F. Addison. (2000). High PCB concentrations in free-ranging Pacific killer whales (Orcinus orca): effects of age, sex and dietary preference.Marine Pollution Bulletin 40: pp. 504-515.

Saulitis, E., C. Matkin, L. Barrett-Lennard, K. Heise and G. Ellis. 2000. Foraging Strategies of Sympatric Killer Whale (Orcinus orca) Populations in Prince William Sound, Alaska. Marine Mammal Science 16: pp.94-109.

Scheffer, V. B. and J. W. Slipp. 1948. The whales and dolphins of Washington State with a key to the cetaceans of the west coast of North America. American Midland Naturalist 39: pp. 257-337.

Schmidt-Nielsen, K. 1997. Animal Physiology: Adaptation and Environment, 5th Ed. Cambridge University Press.Shevchenko, V. I. 1975. Nature of correlations between killer whales and other cetaceans (in Russian). Morskiye Mlekopitayuschchiye 1975: pp. 173-175.

Shpak, O.V. 2012. Mammal-eating killer whales (Orcinus orca) in the western part of The Okhotsk Sea: our observations and interview data. In Materials of the Killer Whale Workshop VII International Conference: Marine Mammals of the Holarctic.Suzdal, Russia. Pp. 17-21.

Shulman-Janiger, A., N. Black and R. Ternullo. Killer Whales of California. 2011. In Whalewatcher Killer Whale: The Top, Top Predator. Pitman, R. L. (Ed.) Journal of the American Cetacean Society. 40(1): 46-47.

Slijper, E.J. 1979. Whales. Ithaca, New York. Cornell University Press.Similä, T. 1997. Behavioral Ecology of Killer Whales in Northern Norway. Dr. Science Thesis, Norwegian College of Fisheries Science, University of Tromsø, Tromsø, Norway.Spencer, M.P., T.A. Gornall and T.C. Poulter. 1967. Respiratory and cardiac activity of killer whales. Journal of Applied Physiology 22, 974-981.

St. Leger, Judy, Guang Wu, M. Anderson, L. Dalton, E. Nilson and D. Wang. 2007. West Nile Virus Infection in Killer Whale, Texas, USA, 2007. Emerging Infectious Diseases (accessed September, 2011). http://www.cdc.gov/EID/content/17/8/101979.htm

Stevens, T. A., D. A. Duffield, E. D. Asper, K. G. Hewlett, A. Bolz, L. J. Gage and G. D. Bossart. 1989. Preliminary findings of restriction fragment differences in mitochondrial DNA among killer whales (Orcinus orca). Canadian Journal Zoology 67:2592–2595.

Sweeney, D. L. 2009. Learning in Human-Dolphin Interactions at Zoological Facilities. PhD Dissertation. University of California, San Diego. 304 pages.

Szymanski, M. D., D. E. Bain, K. Kiehl, S. Pennington, S. Wong and K. R. Henry. 1999. Killer whale (Orcinus orca) hearing: Auditory brainstem response and behavioral audiograms. J. Acoust. Soc. Am. 106: pp.1134-1141.

Tarpy, Cliff. 1979. Killer Whale Attack! National Geographic, April: pp. 542-545.

Taylor, B. L., R. Baird, J. Barlow, S.M. Dawson, J. Ford, J.G. Mead, G. Notarbartolo di Sciara, P. Wade and R. L. Pitman. 2013. Orcinus orca. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.2. www.iucnredlist.org. Downloaded on 20 December 2013.

Tomilin, A. G. 1957. Mammals of the U.S.S.R. and Adjacent Countries. Vol. IX. Cetacea. Moscow, Soviet Union (English translation, 1967, Israel Program for Scientific Translations, Jerusalem).

Thomsen, F., D. Franck and J. K. B. Ford. 2002. On the communicative significance of whistles in wild killer whales (Orcinus orca). Naturwissenshaften 89:404-407.

Titze, I. R. 1994. Principles of Voice Production, Prentice Hall. 354 pages.Tyack, P. L. 1999. Communication and Cognition. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press. Washington, pp. 287-323.

Visser, I.N. 2005. First observations of feeding on thresher (Alopias vulpinus) and hammerhead (Sphyrna zygaena) sharks by killer whales (Orcinus orca), which specialize on Elasmobranchs as prey. Aquatic Mammals 31:83-88.

Visser, I. N. 1999. Benthic foraging on stingrays by killer whales (Orcinus orca) in New Zealand waters. Marine Mammal Science. 15(1): pp. 220-227.

Visser, Ingrid N. 1998. Prolific Body Scars and Collapsing Dorsal Fins on Killer Whales (Orcinus orca) in New Zealand Waters. Aquatic Mammals. 24.2: pp. 71-81.

Visser, I. N., J. Berghan, R. van Meurs and D. Fertl. 2000. Killer whale (Orcinus orca) predation on a shortfin mako shark (Isurus oxyrinchus) in New Zealand waters. Aquatic Mammals 26:229-234.

Visser, I. N. and F. J. Bonoccorso. 2003. New observations and a review of killer whale (Orcinus orca) sightings in Papua New Guinea waters. Aquatic Mammals. 29(1): 150-172.

Visser, I. N., T. G. Smith, I. D. Bullock, G. D. Green, O. G. L. Carlsson and S. Imberti. 2008. Antarctic peninsula killer whales (Orcinus orca) hunt seals and a penguin on floating ice. Marine Mammal Science. 24(1): 225-234.

Walker, L. A., et al. 1988. Urinary concentrations of ovarian steroid hormone metabolites and bioactive FSH in killer whales (Orcinus orca) during ovarian cycles and pregnancy. Biology of Reproduction 39:1013–1020.

Wartzok, D. and D. R. Ketten. 1999. Marine Mammal Sensory Systems. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press. Washington: pp. 117-175.

Wells, R. S., D. J. Boness and G. B. Rathbun. Behavior. In: Reynolds J. E. and S. A. Rommel, Biology of Marine Mammals. Smithsonian Institution Press. Washington. 1999. Pp. 324-422.

White, D., N. Cameron, P. Spong and J. Bradford. 1971. Visual Acuity of the Killer Whale (Orcinus orca). Experimental Neurology 32: pp. 230-236.Whitehead, H. 1985. Why whales leap. Scientific American, 252(3), 84-93.

Wiles, G. J. 2004. Washington State Status Report for the Killer Whale. Washington Department Fish and Wildlife, Olympia. 106 pp.

Williams, T.M. 2009. Swimming. In: Perrin W. F., B. Wursig and J. G. M. Thewissen. Encyclopedia of Marine Mammals, San Diego, CA: Academic Press: pp. 1140-1147.

Williams, R., et al. 2011. Competing conservation objectives for predators and prey: estimating killer whale prey requirements for chinook salmon. PLoS One 6:e26738.

Worthy, Graham A. J., Tamara A. M. Worthy, Pamela K. Yochem and Christopher Dold. 2013. Metabolism of an adult male killer whale (Orcinus orca). DOI: 10.1111/mms.12091.

Ylitalo, Gina M., Craig O. Matkin, Jon Buzitis, Margaret M. Krahn, Linda L. Jones, Teri Rowles and John E. Stein. 2001. Influence of life-history parameters on organochlorine concentrations in free-ranging killer whales (Orcinus orca) from Prince William Sound, AK. The Science of the Total Environment: pp. 183-203.

Yonezawa, M., H. Nakamine, T. Tanaka and T. Miyaji. 1989. Hodgkin's Disease in a Killer Whale (Orcinus orca). Journal of Comparative Pathology. Volume 100, Issue 2: pp. 203-207.


Boston Globe: http://www.bostonglobe.com/news/nation/2014/07/03/some-seaworld-mammals-survive-longer-captivity/JWxo6cwSRpmVXfEW9RRrxK/story.html (accessed April 2015)

CBC News: http://www.cbc.ca/news/canada/north/greenland-hunting-more-killer-whales-as-climate-changes-1.2782297 (accessed May 2015)

CITES http://www.cites.org/eng/app/appendices.php (accessed April 2015)

IUCN Red List of Threatened Species: http://www.iucnredlist.org/details/15421/0

NOAA Killer Whale fact sheet: http://www.nmfs.noaa.gov/pr/species/mammals/cetaceans/killerwhale.htm

NOAA Responsible Marine Wildlife Viewing: http://www.nmfs.noaa.gov/pr/education/viewing.htm (accessed April 2015)

Society for Marine Mammalogy: Committee on Taxonomy. 2012. List of marine mammal species and subspecies. Society for Marine Mammalogy, www.marinemammalscience.org, consulted on May 12, 2014. http://www.marinemammalscience.org/index.php?option=com_content&view=article&id=645&Itemid=340

Southwest Fisheries Science Center, Killer Whale: https://swfsc.noaa.gov/mmtd-killerwhale/Wall Street Journal: http://www.wsj.com/articles/ad-targets-seaworld-over-killer-whales-1405703464 (accessed April 2015)

Photo Credits

Photo Credit: 600x300-8628266356_9d20836694.jpg. Source: Flickr. Image by: Len2040. Year Created: 21 June 2005. Website: www.flickr.com/photos/lenjoh/8628266356/in/album-72157633187981930/ License: CC by ND 2.0. No changes were made to the photo.

Photo Credit: 600x300-1254820034_b7c24ffe0c_b.jpg. Source: Wikimedia Commons. Image by: Josh Larios. Year Created: 27 August 2007. Website: https://commons.wikimedia.org/wiki/File:Chinook_salmon1.jpg License: CC by SA 2.0. No changes were made to the photo.

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Photo Credit: Stranded-killer-whale.jpg. Source: NOAA. Image by: Hawaiian Islands Stranding Response Group. Year Created: N.d.. Website: http://www.fpir.noaa.gov/PRD/prd_marine_mammal_response.html. License: Public Domain. No changes were made to the photo.