References

Ainley, D.G. The Adélie Penguin: Bellwether of Climate Change. New York, Columbia University Press. 2002.

Ainley, D.G., LeResche, R.E. and W.J.L. Sladen. Breeding Biology of the Adélie Penguin. Berkeley. University of California Press. 1983.

Ancel, A., Kooyman, G.L., Ponganis, P.J., Gendner, J.P., Lignon, J., Mestre, X., Huin, N., Thorson, P.H., Robisson, P. and Y. Le Maho.  "Foraging behavior of emperor penguins as a resource detector in winter and summer". Nature. 360:336 – 339. 1992.

Araya, B. and F. Todd. Status of the Humboldt penguin in Chile following the 1982–83 El Niño. Spheniscid Penguin Newsletter. 1(1): 8–10. 1988.

Angelier, F., Barbraud, C., Lormee, H., Prud'homme, F., & Chastel, O. Kidnapping of chicks in emperor penguins: a hormonal by-product? The Journal of Experimental Biology, 1413-1420. 2006.

Australian Antarctic Division, Emperor Penguins: antarctica.gov.au/about-antarctica/wildlife/animals/penguins/emperor-penguins

Baldassin, P., Taniquchi, S., Gallo, H., Silva, R.H. and R.C. Montone. Persistent organic pollutants in juvenile Magellan penguins (Spheniscus magellanicus) found on the northern shore of the state of São Paulo and southern shore of the state of Rio de Janeiro, Brazil. Marine Pollution Bulletin. 64(11): 2502–2506. 2012.

Ballard, G. and David G. Ainley. Short Note killer whales harassment of Adélie penguins at Ross Island. Antarctic Science 17 (#), 385-386. 2005.

Ballard, G., Toniolo, V., Ainley, D.G., Parkinson, C.L., Arrigo, K.R. and P.N. Arrigo. Responding to climate change: Adélie penguins confront astronomical and ocean boundaries. Ecology. 91(7): 2056–2069. 2010.

Banks, J.C., Mitchell, A.D., Waas, J.R. and A.M. Paterson. An unexpected pattern of molecular divergence within the blue penguin (Eudyptula minor) complex. Notornis. 49: 29–38. 2002.

Barham, P.J., Underhill, L.G., Crawford, R.J.M. and T. M. Leshoro. Differences in breeding success between African Penguins (Spheniscus demersus) that were and were not oiled in the MVTreasure oil-spill in 2000. Emu. 107(1): 7–13. 2007.

BBC Earth News. 2009. news.bbc.co.uk/earth/hi/earth_news/newsid_8033000/8033782.stm

Bengtson, J.L., Croll, D.A. and M.E. Goebel. Diving behavior of chinstrap penguins at Seal Island. Antarctic Science. 5(1): 9–15. 1993.

Bingham, Mike. Food Transfer between Gentoo penguin Pygoscelis papua siblings. Marine Ornithology 25:70. 1997.

BirdLife International: birdlife.org

Bohec, LC, Gauthier-Clerc M, Maho LY. The Adaptive Significance of Crèches in the King Penguin. Animal Behaviour. 70:527-538. 2005.

Bowmaker, J.K. and G.R Martin. Visual pigments and oil droplets in the penguin, Spheniscus humbolti. Journal of Comparative Physiology, A. 156: 71–77. 1985.

Brooke, M. and T. Birkhead, Eds. The Cambridge Encyclopedia of Ornithology. New York, Cambridge University Press. 1991.

California Academy of Sciences: calacademy.org/medialibrary/blogs/thelongview/category/dispatches-from-the-voyage/antarctica/cape-royds

Campbell, B. and E. Lack., Eds. Dictionary of Birds. Vermillion, South Dakota, Buteo Books. 1985.

Carroll, R.L. Vertebrate Paleontology and Evolution. New York, W.H. Freeman and Company. 1988.

Cherel, Y., Tremblay, Y., Guinard, E., and Georges, J. Y. Diving behavior of northern Rockhopper Penguin (Eudyptes chrysocome moseleyi) during the brooding period at Amsterdam Island (Southern Indian Ocean). Marine Biology 134:375-385. 1999.

Cherel, Y., Stahl, J.C. and Y. Le Mayo. Ecology and physiology of fasting in king penguin chicks. Auk. 104:254–262. 1987.

Cherel, Y., Robin, J.P., Walch, O., Karmann, H., Netchitailo, P. and Y. Le Maho. Fasting in king penguin. Hormonal and metabolic changes during breeding. American Journal of Physiology. 254: 170–177. 1988.

Chiuchio,A.L., Dickhut, R.M., Cochran, M.A. and H.W. Ducklow. Persistent organic pollutants at the base of the Antarctic marine food web. Environmental Science & Technology. 38(13): 3551–3557. 2004.

Coffin, H.R., Watters, J.V. and J.M. Mateo. Odor-based recognition of familiar and related conspecifics: A first test conducted on captive Humboldt penguins (Spheniscus humboldti). PLoS ONE. 6(9): e25002.doi:10.1371/journal.pone.0025002. 2011.

Collar, N.J. and P. Andrew. Birds to Watch, ICBP Technical Publication No. 8. Snithsonian Press. 1988.

Cool Antarctica: Emperor penguins: coolantarctica.com/Antarctica fact file/wildlife/Penguin_royalty_King_and_Emperor_penguins.htm

Cooper, A. and D. Penny. Mass survival of birds across the Cretaceous-Tertiary boundary: Molecular evidence. Science. 275: 1109-1113. 1997.

Copeland, Karen E. Concentrated small-group foraging behavior in Gentoo penguins Pygoscelis papua. Marine Ornithology 36: 193-194. 2008.

Croxall, J.P. and R.W. Davis. Metabolic rate and foraging behavior of Pygoscelis and Eudyptes penguins at sea. In Penguin Biology: L.S. Davis and J.T. Darby, Eds. San Diego, Academic Press. Pp. 207–228. 1990.

Culik, B.M., Putz, K., Wilson, R.P., Allers, D., Lage, J., Bost, C.A., and Y. Le Maho. Diving energetics in king penguins (Aptenodytes patagonicus). Journal of Experimental Biology. 199: 973-983. 1996.

Cunningham, G.B., Strauss, V. and P.G. Ryan. African penguins (Spheniscus demersus) can detect dimethyl sulfide, a prey-related odour. The Journal of Experimental Biology. 2001: 3123–3127. 2008.

Davis, L. Timing of nest relief and its effect on breeding success in Adélie penguins (Pygoscelis adeliae). Condor. 84: 178–183. 1982.

Davis, L.S. and J.T. Darby, Eds. Penguin Biology. San Diego. Academic Press, Inc. 1990.

Davis, L.S. and E. A.H. Speirs. Mate choice in penguins. In Penguin Biology. Davis, L.S. and J.T. Darby, Eds. San Diego. Academic Press, Inc. 1990.

del Hoyo, J., Elliot, A., and J. Sardatal, eds. Handbook of Birds of the World, Vol. 1. Ostrich to Ducks. Barcelona, Lynx Editions. 1992.

Despin, B. Oiseau Revue fr. Orn. 42:69–83. 1972. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Dresp, B. Jouventin, P. Langley, K. Ultraviolet reflecting photonic microstructures in the King Penguin beak. Biology Letters. Volume 1, Number 3, 310-313. 2005.

Ellis, S. The penguin conservation management plan (CAMP): Introduction and Overview. Penguin Conservation. June:13–17. 1993.

Falla, R.A. Rep. B.A.N.Z. Antarc. Res. Exp. Ser. B., II. 1937. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Froget, G., Butler, P.J., Woakes, A.J., Fahlman, A., Kuntz, G., Le Maho, Y. and Y. Handrich. Heart rate and energetic of free-ranging king penguins (Aptenodytes patagonicus ). The Journal of Experimental Biology. 207: 3917–3926. 2004.

Frost, P.G.H., Siegfried, W.R., A.E. Burger. Behavioural adaptations of the jackass penguin, Spheniscus demersus to a hot arid environment. Journal of Zoology. 179(2):165–187. 1976.

Geisz, H.N., Dickhut, R.M., Cochran, M.H., Fraser, W.R. and H.W. Ducklow. Melting Glaciers: A Probable Source of DDT to the Antarctic Marine Ecosystem. Environmental Science & Technology. 42:3958–3962. 2008.

GPS (Global Penguin Society): globalpenguinsociety.org/about-us.html

Groscolas, R. Metabolic adaptations to fasting in emperor and king penguins. In Penguin Biology, Marchant and J.T Darby, Eds. San Diego, Academic press. 269–296. 1990.

Harrington, H.J. Nortornis. 9:33–39. 1960. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Harrison, G.L., McLenachan, P.A., Phillips, M.J., Slack, K.E., Cooper, A., and D. Penny. Four new avian mitochondrial genomes help get to basic evolutionary questions in the late Cretaceous. Molecular Biology and Evolution. 21(6): 974-983. 2004.

Howland, H.C. and J.G. Sivak. Penguin vision in air and water. Vision Research. 24(12): 1905-1909. 1984.

International Bird Rescue: bird-rescue.org

International Penguin Conservation Work Group (IPCWG): penguins.cl/about-penguins.htm

Johnstone, G.W. ANARE Scient. Rep. Ser. B, 1. 123:1–62. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Jouventin, P., Barbraud, C. and Rubin, M. Adoption in the emperor penguin. Animal Behaviour. 50,1023 -1029. 1995.

Kooyman, G.L., Drabek, C.M., Elsner, R., and W.B. Campbell. Diving behavior of the emperor penguin Aptenodytes forsteri. Auk. 88:775-795. 1971.

Kooyman, G.L., Cherel, Y., Le Maho, Y., Croxall, J.P., Thorson, P.H., Ridoux, V. and C.A. Kooyman. Diving behavior and energetics during foraging cycles in king penguins. Ecological Monographs. 62(1): 143-163. 1992.

Kooyman, G.L., Ponganis, P.J., Castellini, M.A., Ponganis, E.P., Ponganis, K.V.,Thorson, P.H., Eckert, S.A., and Y. LeMaho. Heart rates and swim speeds of emperor penguins diving under sea ice. Journal of Experimental Biology. 165(1): 161-180. 1992.

Kooyman, G.L. Breeding habits of emperor penguins in the western Ross Sea. Antarctic Science. 5(2): 143-148. 1993.

Kooyman, G.L. and T.G. Kooyman. Diving behavior of emperor penguins nurturing chicks at Coulman Island, Antarctica. The Condor. 97:536-549. 1995.

Ksepka, D.T., Bertelli, S. and N.P. Giannini. The phylogeny of the living and fossil Sphenisciformes (penguins). Cladistics. 22 (5): 412–441. 2006.

Lamey, T. Hatch asynchrony and brood reduction in penguins. In Penguin Biology: L.S. Davis and J.T. Darby, Eds. San Diego, Academic Press. Pp. 207–228. 1990.

Marchant, S. and P.J. Higgins, Eds . Handbook of Australian, New Zealand, and Antarctic Birds. Melbourne: Oxford University Press. 1990.

Martin, G.R. Eye structure and foraging in king penguins Aptenodyte patagonicus. Ibis. 141 (3): 444–450. 2008.

Martin, G.R. and S.R. Young. The eye of the Humboldt penguin, Speniscus humbolti: Visual fields and schematic optics. Proceedings of the Royal Society of London, B. 223: 197–222. 1984.

McGonigal, David. Antarctica. Secrets of the Southern Continent. Australia: Firefly Books, Ltd. 2008.

Meir, J.U., Stockard, T.K., Williams, C.L., Ponganis, K.V. and P.J. Ponganis. Heart rate regulations and extreme bradycardia in diving emperor penguins. The Journal of Experimental Biology. 211: 1169–1179. 2008.

Mills, Kyra L. Multispecies Seabird Feeding Flocks in the Galápagos Islands. The Condor

Vol. 100, No. 2, pp. 277-285. 1998.

Müller-Scwarze, D. The Behavior of Penguins: Adapted to Ice and Tropics. Albany, New York. State University of New York Press. 1984.

Murphy, R.C. Auk. 64:454–455. 1947. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Nel, D.C. and P.A. Whittington, Eds. Rehabilitation of Oiled African Penguins: A Conservation Success Story. Cape town: BirdLife South Africa and the Avian Demography Unit. University of Cape Town. 2003.

Penguin Project: mesh.biology.washington.edu/penguinProject

Penguin Net: penguin.net.nz/species/yep

Penguin Science: penguinscience.com

Penguin TAG. Spheniscus Penguin Husbandry Manual, First Ed. EAZA. 1993.

Penguin TAG (Penguin Taxon Advisory Group): zoopenguins.org

Penguin World: penguinworld.com

Perkins, S. Flightless, feathered friends: New tales of penguin evolution, past and present. Science News. 166(22): 346–348. 2004.

Peterson, R.T. Penguins. Boston, Houghton Mifflin Company. 1979.

Polito, M.J. and W.Z. Trivelpiece. Transition to independence and evidence of extended parental care in the gentoo penguin (Pygoscelis papua). Marine Biology. 154: 231–240. 2008.

Ponganis, P.J. and G. L. Kooyman. Diving physiology of penguins. Acta XX Congressus Internationalis Ornithologici. Wellington, New Zealand Ornithological Congress Trust Board. 1887–1891. 1990.

Ponganis, P.J., Kooyman, G.L., Van Dam, R., and Y. LeMaho. Physiological responses of king penguins during simulated diving to 136 m depth. Journal of Experimental Biology. 202:2819-2822. 1999.

Ponganis, P.J., Stockard, T.K., Meir, J.U., Williams, C.L., Ponganis, K.V., van Dam, R.P. and R. Howard. Returning on empty: extreme blood O2 depletion underlies dive capacity of emperor penguins. Journal of Experimental Biology. 210: 4279–4285. 2007.

Ponganis, P.J., Van Dam, R.P., Knower, T. and D.H. Levenson. Temperature regulation in emperor penguins foraging under sea ice. Comparative Biochemistry and Physiology: Part A. 129:811-820. 2001.

Ponganis, P.J., Van Dam, R.P., Levenson, D.H. , Knower, T., Ponganis, K.V., and G. Marshall. Regional heterothermy and conservation of core temperature in emperor penguins diving under sea ice. Comparative Biochemistry and Physiology: Part A. 135:477-487. 2003.

Ponganis, P.J., Van Dam, R.P., Knower, T., Levenson, D.H. , and K.V. Ponganis, Deep dives and aortic temperatures of emperor penguins: new directions for bio-logging at the isolated dive hole. Memoirs of the National Institute of Polar Research (Special Issue). 58:155-161. 2004.

Prévost, J. and V. Vilter. Histologie de la secrétion oesophagienne du Manchot empereur. In Proceedings of the 13th International Ornithological Congress. 1085–1094. 1963.

Pütz, K. and Y. Cherel. The diving behaviour of brooding king penguins ( Aptenodytes patagonicus) from the Falkland Islands: variation in dive profiles and synchronous underwater swimming provide new insights into their foraging strategies. Marine Biology. 147: 281–290. 2005.

Quintana, R.D. Nest-site characteristics of a Gentoo Penguin Pygoscelis papua colony at Cierva Point, Antarctic Peninsula. Marine Ornithology 29: 107–110. 2001.

Rand, R.W. Ibis. 96:171–206. 1954. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Ritter, L., Soloman, K.R., Forget, J., Stemeroff, M. and C. O'Leary. A review of selected Persistent Organic Pollutants: DDT-Aldrin-Dieldrin-Endrin-Chlordane, Heptachlor-Hexachlorobenzene-Mïrex-Toxaphene, Polychlorinated biphenyls, Dioxins and Furans. The International Programme on Chemical Safety (IPCS) within the framework of the Inter-Organization Programme for the Sound Management of Chemicals (IOMC). 1995.

Robertson, G. Australian Wild. Res. 13: 583–587. 1986. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Robertson, G.G. The foraging ecology of emperor penguins Aptenodytes forsteri at two Mawson Coast colonies, Antarctica. ANARE Report 138. 139 pp. 1995.

Sato, K., Naito, Y., Kato, A., Niizuma, Y., Watanuki, Y., Charrassin, J.B., Bost, C.A., Handrich, Y., and T. Le Maho. Buoyancy and maximal diving depth in penguins: do they control inhaling air volume? Journal of Experimental Biology. 205: 1189-1197. 2002.

Sato, K., Shiomi, K., Marshall, G., Kooyman, G.L. and P.J. Ponganis. Stroke rates and diving air volumes of emperor penguins: implications for dive performance. Journal Experimental Biology. 214: 2854–2863. 2011.

Scholander, P.F. Hvalradets Skrifter, 1940. In Kooyman, G.L., Ponganis, P.J., Castellini, M.A., Ponganis, E.P., Ponganis, K.V.,Thorson, P.H., Eckert, S.A., and Y. LeMaho. Heart rates and swim speeds of emperor penguins diving under sea ice. Journal of Experimental Biology. 165(1): 161-180. 1992.

Scientific Committee on Antarctic Research (SCAR). Persistent Organic Pollutants (POPs) in the Antarctic environment: A Review of Findings. The SCAR Action Group on Environmental Contamination in Antarctica. February, 2009.scar.org/publications/occasionals/POPs_in_Antarctica.pdf

Scolaro. J.A. Hall, M., Ximinez, I. and O. Kovacs. El pinguino de Magellanes (Spheniscus magellanicus): Biologia y desarrollo de la incubacion en la colonia de Punto Tombo, Chubut, Argentina. Ecologia (Museum Argentino de Ciencias Natruales, Buenos Aires). II (5): 103–110. 1980.

Simpson, G.G. Penguins: Past and Present, Here and There. New Haven, Connecticut. Yale University Press. 1976.

Slack, K.E., Jones, C.M., Ando, T., Harrison, G.L., Fordyce, R.E., Arnason, U. and D. Penny. Early penguin fossils, plus mitochondrial genomes, calibrate avian evolution. Molecular Biology and Evolution. 23 (6): 1144–1155. 2006.

Sparks, J. and T. Soper. Penguins. New York. Facts On File Publications. 1987.

Stahl, J.C. 4th Symposium Antarctic Biology. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds. Melbourne: Oxford University Press. 1990.

Steinfurth, Antje. Marine ecology and conservation of the Galápagos penguin, Spheniscus mendiculus. Report to the Charles Darwin Research Station and the Galápagos National Park Service, Puerto Ayora, Isla Santa Cruz, Galápagos, Ecuador. 2007.

Takahashi A., K. Sato, J. Nishikawa, Y. Watanuki and Y. Naito. Synchronous diving behavior of Adelie penguins. Journal of Ethology. DOI 10.1007/s10164-003-0111-1. 2003.

Tremblay Y. and Y. Cherel. Benthic and pelagic dives: a new foraging behaviour in rockhopper penguins. Marine Ecology Prog Ser. 204:257–267. 2000.

Stonehouse, B., Ed. The Biology of Penguins. Baltimore, University Park Press. 1975.

Trivelpiece, W.Z., Bengston, J.L., Trivelpiece, S.G. and N.J. Volkman. Foraging behavior of gentoo and chinstrap penguins as determined by new radiotelemetry techniques. The Auk . 103: 777–781. 1986.

Trivelpiece, W.Z., Hinke, J.T.,Miller, A.K., Reiss, C.S., Trivelpiece, S,G. and George M. Watters. Variability in krill biomass links harvesting and climate warming to penguin population changes in Antarctica. PNAS. 108 (18). 2011.

Trivelpiece, W. and S. Trivelpiece. Courtship period of Adélie, gentoo, and chinstrap penguins. In Penguin Biology, Marchant and J.T Darby, Eds. San Diego, Academic press. 269–296. 1990.

Villiers, de M. S. and P. J. N. de Bruyn. Stone-swallowing by three species of penguins. Marine Ornithology 32: 185–186. 2004.

Welty, J.C. The Life of Birds, 3rd Ed. Philadelphia, Saunders College Pub. 1982.

Wever, E.G., Herman, P.N., Simmons, J.A. and D.R. Hertzler. Hearing in the black-footed penguin, Spheniscus demersus, as represented by the cochlear potentials. Proc. Natl. Acad. Sci. U.S.A. 63(3): 676–680. 1969.

White, M.G. and J.W.H. Conroy. Ibis. 118: 371–373. 1975. In Handbook of Australian, New Zealand and Antarctic Birds. Marchant, S. and P.J. Higgins, Eds.